back

Key of European species of Tomentella incl. Odontia 

 

 

-        spores size without ornaments

-        cyanescent-  hyphae and basidia bluish-green to bluish black in KOH

 

 

 

 

 

A- Hyphae clamped

1

 

B- Clamps absent or occasionally present

38

 

 

 

 

1a.   Cystidia present

2

 

1b.   Cystidia absent

8

 

 

 

 

2a.   Cystidia hyphoid 55-95 x 3-4 m, growing from subiculum and rhizomorphs,

 

 

        hymenophore hazel-brown to olive-brown, smooth to granulose,

 

 

        margin byssoid with brownish dimitic rhizomorphs,

 

 

        spores irregular 8,5-10 m 

 

 

        Tomentella atroarenicolor Nikol. 1970

 

 

 

 

 

2b.   Cystidia acuminate, capitate or clavate

3

 

 

 

 

3a.   Cystidia acuminate (basally widened, rarely hyphoid)

4

 

3b.   Cystidia capitate

5

 

3c.   Cystidia clavate

6

 

 

 

 

4a.   Hymenophore  hazel-brown, granulose,

 

 

        cystidia somewhat distended base and tapering towards the slightly,

        expanded apices, often apical encrusted with brown deposits,

 

 

        some spores reddish brown in KOH

 

 

        Tomentella subtestacea Bourd. & Galz.1924

 

 

 

 

 

4b.   Hymenophore greyish-green, smooth to granulose, mucedinoid,

 

 

        cystidia apical slightly incrusted

 

 

        Tomentella galzinii Bourd. 1924

 

 

 

 

 

5a.   Cystidia pale brownish to greenish

 

 

        basidiocarps separable, dimitic rhizomorphs present,

 

 

        hymenophore yellowish-brown to olive-brown,

 

 

        Tomentella pilosa (Burt) Bourd. & Galz. 1924

 

 

 

 

 

5b.   Cystidia hyaline,

 

 

        basidiocarps adherent, without rhizomorphs,

 

 

        hymenophore greyish-green to olive-brown,

 

 

        Tomentella viridula Bourd.& Galz.1924

 

 

 

 

 

6a.   Basidiocarps separable, rhizomorphs present,                             

 

 

        hymenophore greyish-brown to hazel-brown, smooth,
        subicular hyphae brownish, thick-walled

 

        Tomentella subpilosa Litsch 1960

 

 

        (similar to T. muricata, an American species)

 

 

 

 

 

6b.   Basidiocarps adherent, rhizomorphs absent,

 

 

        subicular hyphae hyaline and thin-walled                       

7

 

 

 

7a.   Spores irregular, partly lobed

 

 

        Tomentella clavigera Litsch. 1960

 

 

 

 

 

7b.   Spores regular globose to ellipsoid,

 

 

        subiculum thin to absent,

 

 

        subhymenial hyphae partly short celled and inflated

 

 

        Tomentella subclavigera Litsch. 1933

 

 

 

 

 

8a.   Spores average < 6,5m 

9

 

8b.   Spores average > 6,5 m

13

 

 

 

 

9a.   Spores 3-4 m,

 

 

        hymenophore with short spines, rusty yellowish

 

 

        Tomentella (Odontia) duemmeri (Wakef.) M.J. Larsen 1974

 

 

 

 

 

9b. Spores average 4,5-6,5 m 

10

 

 

 

 

10a.  Spores tubercular, hymenophore ferruginous-brown,

 

 

        smooth to hydnoid, margin byssoid, rhizomorphs dimitic

 

 

        Tomentella (Odontia) calcicola (Bourd. & Galz.) M.J. Larsen 1967

 

 

 

 

10b.  Spores echinulate

11

 

 

 

 

11a.  Basidiocarps adherent, rhizomorphs absent, hymenophore greyish brown

 

 

basidia up to 25 m long

 

 

Tomentella griseoumbrina Litsch. apud Lundell & Nannf. 1936  

 

 

 

 

 

11b. Basidiocarps separable, rhizomorphs present

12

 

 

 

 

12a. Hymenophore rosy brown, smooth

 

 

typical yellowish rhizomorphs present

 

 

Tomentella palliodocastanea X. Lu, Y.H. Mu & H.S. Yuan 2018

 

 

 

 

 

12b. Hymenophore greyish, granulose, rhizomorphs white

 

 

Tomentella cinerascens (P. Karst.) Hoehn. & Litsch. 1906

 

 

 

 

 

13a.  Spore ornaments bifurcate, Hymenophore hydnoid

14

 

13b.  Spore ornaments not bifurcate or with a view of bifurcate warts                              

 

Hymenophore smooth to granulare

15

 

 

14a.  Spores brown, slightly globose 7-8 m, with short pines,

 

 

         rhizomorphs dimitic

 

 

         Tomentella (Odontia) crinalis (Fr.) M.J. Larsen 1967        

 

 

 

 

 

14b.  Spores hyaline, irregular subglobose, 8-10 m

 

         rhizomorphs monomitic

 

 

         Tomentella italica (Sacc.) M.J. Larsen 1967

 

 

 

15a.  Immaturity basidia vesicular up to 25 m wide, 

 

 

         hymenophore smooth, fawn, basidiocarps compact,

 

 

         spores irregular subglobose 7-9,5 m

 

 

         Tomentella terrestris (Berk. & Broome) M.J. Larsen 1974

 

 

 

 

 

15b.  Immaturity basidia clavat to utriform, < 15 m wide                            

16

 

 

 

 

16a.  Spores average 6-7,5 m (if hyphae short celled and inflated

17

 

(if hyphae partly short celled and inflated go to 32a)

 

 

16b.  Spores average larger

19

 

 

 

 

17a.  Subicular hyphae thick-walled, brown, incrusted, up to 6 m wide,

 

 

         hymenophore greyish-brown to dark brown, granulose,

 

 

         spores partly irregular globose, 6-8 m

 

         Tomentella neobourdotii M.J. Larsen 1968

 

 

 

 

17b.  Subicular hyphae thin-walled, hyaline to pale brown 

18

 

 

18a.  Basidia and hyphae strong cyanescent, hymenophore blackish brown,

 

 

         rhizomorphs distinct, margin and subiculum ferruginous-yellowish

 

 

         Tomentella botryoides (Schwein.) Bourd. & Galz. 1924

 

(if rhizomorphs dimitic and hymenophore olivaceous brown

 

 

comp. T. ferruginea 36a.)

 

 

 

18b.  Basidia and hyphae not cyanescent, rhizomorphs absent

 

 

         hymenophore brownish, margin and subiculum same coloured, 

 

 

spores not bifurcate

 

 

         Tomentella sp.  FD 7548

 

 

(?Tomentella microspora (P. Karst.) Hhn. & Litsch. 1906 ss. Ales Jirsa)

 

 

 

19a. Hymenophore whitish, yellowish or pale ochraceous to mustard yellow

        Tomentella ochraceo-olivacea Litsch. 1933  

        (Syn. Tomentella fatrensis Svrček 1958)

 

 

 

 

 

 

19b. Hymenophor pale to dark brown, olivaceous or rusty brown

20

 

 

 

 

20a.  Spores subglobose or ellipsoid, not lobed          

21

 

20b.  Spores with irregular outline, mostly lobed     

30

 

 

 

 

21a.  Spores ellipsoid (lateral view)                     

22

 

21b.  Spores subglobose (lateral view)                        

23

 

 

 

 

22a.  Subicular hyphae hyaline to pale brownish, thin-walled,

 

 

         subhymenial hyphae short celled and inflated,

 

 

         spores 7-10 m long

 

 

         Tomentella ellisii (Sacc.) Jlich & Stalpers 1980

 

 

 

 

 

22b.  Subicular hyphae brown, thick-walled,

 

 

         subhymenial hyphae not short celled and inflated,

 

 

         spores 8-12 m long, partly irregular 

 

 

         Tomentella atramentaria Rostr. 1894

 

 

 

23a.  Subicular hyphae hyaline

24

 

23b.  Subicular hyphae brown

25

 

 

 

 

24a.  Hymenophore greyish-brown, tubercular, spores 8-11 m,

 

 

         basidiocarps separable, rhizomorphs present,

 

 

         subiculum white, margin yellowish to white

 

 

         Tomentella asperula (P. Karst.) Hoehn. & Litsch. 1906

 

 

 

24b.  Hymenophore pale brown to brick, spores 7-8,5 m

 

 

         (forms of T. lateritia 34a. and T. coerulea 35b. with more regular spores)

 

 

 

 

23a.  Hymenophor rust-yellowish, spores yellowish-brown

26

 

23b.  Hymenophor bright to dark brown or olive-greenish, spores brown

27

 

 

26a.  Spores with long spines, globose 8-11 m,

         apiculus amyloid

        Tomentella bryophila (Pers.) M.J. Larsen 1974

 

 

 

26b. Spores with short spines, irregular subglobose up to 6-9 m,

        apiculus not amyloid

        Tomentella ferruginella Bourd. & Galz. 1924

 

 

 

27a.  Hymenophore olive-greenish

         Tomentella viridescens (Bres. & Torr.) Bourd. & Galz. 1928 

 

 

 

 

27b.  Hymenophor bright to dark brown

28

 

 

 

 

28a.  Spores globose, 8 -11 m, spines up to 3 m,

 

 

         Subicular hyphae not incrusted

 

 

         Tomentella stuposa (Link) Stalpers 1984 s. l.

-- Spores 6 - 8 m Tomentella spec.

--      Spores 8 - 10 m  Tomentella stuposa (Link) Stalpers 1984 s. str.

--      Spores 10 - 13 m Tomentella bresadolae (Brinkmann) Hhn. & Litsch. 1908

 

 

 

28b.  Spores subglobose to ellipsoid, 7,5-9,5 m, partly irregular

29

 

 

 

 

29a.  Subicular hyphae incrusted, apiculus not amyloid, with spines up to 2 m

 

 

         Tomentella lapida (Pers.) Stalpers 1984

 

 

         (similar T. fuliginea with amyloid apiculus and subicular hyphae not incrusted)

 

29b.  Subicular hyphae not incrusted, with short spines, apiculus not amyloid

         Tomentella alpina Peintner & Dmmrich 2011

 

 

         (alpine species on Salix herbacea)

 

 

 

 

 

30a.  Spores with long spines up to 2,5 m, hymenophore smooth, fawn to rosy-buff,

 

 

         margin byssoid to fimbriat, rhizomorphs present

 

 

         Tomentella testaceogilva Bourd. & Galz. 1924

 

 

 

 

 

30b.  Not with the above combination of characters

31

 

 

 

 

31a.  Subhymenial hyphae to 8 (10) m wide, short celled and inflated

32

 

31b.  Subhymenial hyphae to 5,5 m wide, regular       

33

 

 

 

 

32a.  Spores rough tuberculate-echinulate,       

         a few warts bifurcate, mostly spores with oil drops, 

         margin often fimbriat, partly rhizomorphs present

 

 

         Tomentella radiosa(P. Karst.) Rick 1934

 

 

         (Syn. Thelephora terrestris f. resupinata Donk 1933)

 

 

 

 

 

32b.  Spores with long spines 7-9 m,

         hyphae to 10 m wide, inflated up to 15 m wide,

 

 

         margin undetermined, rhizomorphs absent

 

 

         Tomentella sublilacina agg.

 

 

 

 

 

33a.  Subiculum thin, rhizomorphs rarely, 

         basidia partly with reddish contents (examines in distilled water!)

 

 

         hymenial with amber-tinted adhering material, solving in KOH

         and partly cyanescent 

34

 

 

 

 

33b.  Subiculum and rhizomorphs distinctly,    

 

 

         basidia not reddish in distilled water

36

 

 

 

 

34a.  Hymenophore brick-red, not cyanescent

        Tomentella lateritia Pat. 1894

 

 

 

 

34b. Hymenophore brownish

35

 

 

 

 

35a.  Hymeniophore dark brown, distinctly cyanescent            

 

 

         Tomentella molybdaea Bourdot & Galzin 1924   

 

35b.  Hymenophore pale brownish, not cyanescent

 

 

         Tomentella coerulea (Bres.) Hoehn. & Litsch. 1908 ss. M.J. Larsen

 

 

 

 

 

36a.  Basidia and hyphae cyanescent,

 

 

         hymenophore ferruginous-brown to olive-brown, granulose, 

 

 

         margin partly yellowish, rhizomorphs dimitic

 

 

         Tomentella ferruginea (Pers.) Pat. 1887

 

 

         (Remarks: If hymenophore blackish-brown view Tomentella botryoides 19a.)

 

 

 

 

 

36b.  Basidia and hyphae not cyanescent

37

 

   

 

 

37a.  Hymenophore pale brownish with greenish or reddish tints, granulose, margin,

         byssoid, ferruginous-brown, rhizomorphs monomitic or rarely dimitic

 

 

         Tomentella punicea (Alb. & Schwein.) J. Schrt. 1889   

 

 

 

 

 

37b.  Hymenophore ferruginous-reddish to chestnut-brown, smooth to tubercular, 

 

 

         margin ferruginous-reddish, rhizomorphs dimitic

 

 

         Tomentella umbrinospora M.J. Larsen 1968

 

 

 

 

 

38a.  Spores ornaments bifurcate                                       

39

 

38b.  Spores ornaments not bifurcate                           

40

 

 

 

 

39a.  Hymenophore tubercular to hydnoid, cinnamon brown,

 

 

         spores slightly globose with short bifurcate ornaments 6-8 m,

 

 

         with thick-walled up to 200 m long tramal pseudocystidia

 

 

         Tomentella (Odontia) fibrosa (Berk. & Curtis) Kljalg 1996

 

 

 

 

 

39b.  Hymenophore smooth

 

 

         (cf. Pseudotomentella!)

 

 

 

 

 

40a.  Spores brown                                                            

41

 

40b.  Spores hyaline (cf. Tomentellopsis!)

 

 

 

 

 

41a.  Occasionally clamps present,

 

         subicular hyphae and subhymenial hyphae occasionally clamps present,

         spores regular to irregular ellipsoid, 8-12 m long, spines up to 1,5 m

         Tomentella cinereoumbrina (Bres.) Stalpers 1993

 

 

 

 

 

41b.  All hyphae without clamps

42

 

 

 

 

42a.  Spores regular globose 79 m,  spines up to 3 m

 

 

         Tomentella spinosispora Cizek  2004

 

 

         (the type is described with clamps-bearing subicular hyphae,

          but now recognised as alien hyphae, E. Martini 2021)

 

 

 

 

 

42b.  Spores ellipsoid or irregular globose

43

 

 

 

 

43a.  Basidiocarps thick, crustose, mostly on soil

 

 

         Tomentella fuscocinerea (Pers.) Donk 1933

 

 

         (= Thelephora fuscocinerea Pers.)

 

 

 

 

 

43b.  Basidiocarps thin, tomentelloid, on wood

44

 

 

 

 

44a.  Spores with spines up to 2,5 m, spores irregular globose,

 

 

         basidiocarps mucedinoid

 

 

         Tomentella badia (Link) Stalpers 1975

 

 

         (if spores regular globose view 42a.)

 

 

 

44b.  Spores with spines up to 1,5 m, spores regular ellipsoid

 

 

         Tomentella cf. brunneofirma M.J. Larsen 1967