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Key of European species of Tomentella incl. Odontia 

 

 

 

A.    Hyphae clamped

1

B.    Clamps absent or occasionally present

36

 

 

1a.   Cystidia present

2

1b.   Cystidia absent

8

 

 

2a.   Cystidia hyphoid 55-95 x 3-4 m, growing from subiculum and rhizomorphs,

 

        hymenophore hazel-brown to olive-brown, smooth to granulose,

 

        margin byssoid with brownish dimitic rhizomorphs,

 

        spores irregular 8,5-10 m 

 

        Tomentella atroarenicolor Nikol. 1970

 

 

 

2b.   Cystidia acuminate, capitate or clavate

3

 

 

3a.   Cystidia acuminate (basally widened, rarely hyphoid)

4

3b.   Cystidia capitate

5

3c.   Cystidia clavate

6

 

 

4a.   Hymenophore  hazel-brown, granulose,

 

        cystidia somewhat distended base and tapering towards the slightly,

        expanded apices, often apical encrusted with brown deposits,

 

        some spores reddish brown in KOH

 

        Tomentella subtestacea Bourd. & Galz.1924

 

 

 

4b.   Hymenophore greyish-green, smooth to granulose, mucedinoid,

 

        cystidia apical slightly incrusted

 

        Tomentella galzinii Bourd. 1924

 

 

 

5a.   Cystidia pale brownish to greenish

 

        basidiocarps separable, dimitic rhizomorphs present,

 

        hymenophore yellowish-brown to olive-brown,

 

        Tomentella pilosa (Burt) Bourd. & Galz. 1924

 

 

 

5b.   Cystidia hyaline,

 

        basidiocarps adherent, without rhizomorphs,

 

        hymenophore greyish-green to olive-brown,

 

        Tomentella viridula Bourd.& Galz.1924

 

 

 

6a.   Basidiocarps separable, rhizomorphs present,                             

 

        hymenophore greyish-brown to hazel-brown, smooth,
        subicular hyphae brownish, thick-walled

        Tomentella subpilosa Litsch 1960

 

        (similar to T. muricata, an American species)

 

 

 

6b.   Basidiocarps adherent, rhizomorphs absent,

 

        subicular hyphae hyaline and thin-walled                       

7

 

 

7a.   Spores irregular, partly lobed

 

        Tomentella clavigera Litsch. 1960

 

 

 

7b.   Spores regular globose to ellipsoid,

 

        subiculum thin to absent,

 

        subhymenial hyphae partly short celled and inflated

 

        Tomentella subclavigera Litsch. 1933

 

 

 

8a.   Spores up to 6 m  (measurement without spines)                               

9

8b.   Spores larger

10

 

 

9a. Spores 3-4 m,

 

hymenophore short-spined, rusty yellowish

 

Tomentella (Odontia) duemmeri (Wakef.) M.J. Larsen 1974

 

 

 

9b.   Spores tubercular, globose 5-6 m,

 

        hymenophore ferruginous-brown, smooth to hydnoid,

 

        margin byssoid, dimitic rhizomorphs

 

        Tomentella (Odontia) calcicola (Bourd. & Galz.) M.J. Larsen 1967

 

 

9c.   Spores echinulate, globose 4,5-5,5 m, Basidia to 30 m long,

 

        hymenophore brownish, smooth,

 

        without rhizomorphs

 

        Tomentella griseoumbrina Litsch. 1936

 

 

 

10a.  Spores ornaments bifurcate

11

10b.  Spores ornaments not bifurcate                               

12

 

11a.  Hymenophore hydnoid, ferruginous brown,

 

         rhizomorphs dimitic,

 

         spores brownish, subglobose 7-8 m

 

         Tomentella (Odontia) crinalis (Fr.) M.J. Larsen 1967

 

 

 

11b.  Hymenophore tubercular or hydnoid, rare smooth, greyish-brown,

         rhizomorphs monomitic,

 

         spores hyaline, subglobose, partly irregular 8-10 m

 

         Tomentella italica (Sacc.) M.J. Larsen 1967

 

 

 

11c. Hymenophore smooth, ferruginous-brown, without rhizomorphs,

Tomentella sp. FD 8792

 

 

12a.  Immaturity basidia vesicular up to 20 m wide, mature basidia up to 85 m long, 

 

         hymenophore smooth, fawn, basidiocarps compact,

 

         spores irregular subglobose 7-9,5 m

 

         Tomentella terrestris (Berk. & Broome) M.J. Larsen 1974

 

 

 

12b.  Not with this combination of characters                             

13

 

 

13a.  Spores average 6-7 m

        (if hyphae short celled and inflated go to 30)

14

 

 

13b.  Spores average larger

18

 

14a.  Subicular hyphae thick-walled, brown, incrusted, up to 6 m wide,

 

         hymenophore greyish-brown to dark brown, granulose,

 

         spores partly irregular globose

         Tomentella neobourdotii M.J. Larsen 1968

 

 

14b.  Subicular hyphae thin-walled

15

 

 

15a. Hymenophor greyish, brownish or ferruginous-yellowish

16

 

 

15b. Hymenophore whitish to yellowish or pale ochre

Tomentella ochraceo-olivacea Litsch. 1933

(Syn. Tomentella fatrensis Svrček 1958)

 

        

16a.   Hymenophore greyish, smooth to granulose,

          spores globose to subglobose    

          Tomentella cinerascens (P. Karst.) Hoehn. & Litsch. 1906

 

 

 

16b.  Hymenophore brownish to ferruginous-brown, spores irregular, lobed

17

 

17a.  Basidia and hyphae strong cyanescent (hyphae and basidia bluish-green,

         to bluish black in KOH), hymenophore blackish brown,

 

         rhizomorphs distinct, margin and subiculum ferruginous-yellowish

 

         Tomentella botryoides (Schwein.) Bourd. & Galz. 1924

 

17b.  Basidia and hyphae not cyanescent, rhizomorphs absent

 

         hymenophore brownish, margin and subiculum same coloured 

 

         Tomentella sp.  FD 7548

 

 

 

18a.  Spores subglobose or ellipsoid, not lobed          

19

18b.  Spores with irregular outline, often lobed     

28

 

 

19a.  Spores ellipsoid (lateral view)                     

20

19b.  Spores subglobose (lateral view)                        

21

 

 

20a.  Subicular hyphae hyaline, thin-walled,

 

         subhymenial hyphae short celled and inflated,

 

         spores 7-9(10) m long

 

         Tomentella ellisii (Sacc.) Jlich & Stalpers 1980

 

 

 

20b.  Subicular hyphae brown, thick-walled,

 

         subhymenial hyphae not short celled and inflated,

 

         spores 8-12 m long, partly irregular 

 

         Tomentella atramentaria Rostr. 1894

 

 

21a.  Subicular hyphae thin-walled

22

21b.  Subicular hyphae thick-walled

23

 

 

22a.  Hymenophore greyish-brown, tubercular, spores 8-11 m,

 

         basidiocarps separable, rhizomorphs present,

 

         subiculum white, margin yellow to white

 

         Tomentella asperula (P. Karst.) Hoehn. & Litsch. 1906

 

 

22b.  Hymenophore pale brown to brick, spores 7-8,5 m

 

         (forms of T. lateritia 32a. and T. coerulea 33b. with more regular spores)

 

 

23a.  Spores yellowish

24

23b.  Spores brown

26

 

 

24a.  Hymenophore olive-greenish

         Tomentella viridescens (Bres. & Torr.) Bourd. & Galz. 1928 

 

24b. Hymenophor ferruginous-brown

 

25a. Spores with long spines, regular globose 7-11 m,

apiculus amyloid

Tomentella byrophila (Pers.) M.J. Larsen 1974

25

 

25b. Spores with short spines, irregular globose to ellipsoid -8,5 m,

apiculus not amyloid

Tomentella ferruginella Bourd. & Galz. 1924

 

26a.  Spores globose to subglobose, 8,5 -11 m, spines up to 3 m,

 

Subicular hyphae not incrusted

 

         Tomentella stuposa (Link) Stalpers 1984 s. l.

 

-- Spores 6,5 - 8 m cf. Tomentella globosa X. Lu, K. Steffen & H.S. Yuan 2018

-- Spores 8 - 10 m Tomentella stuposa (Link) Stalpers 1984 s. str.

-- Spores 10 - 13 m Tomentella bresadolae (Brinkmann) Hhn. & Litsch. 1908

 

 

26b. Spores subglobose to ellipsoid, 7,5-9, 5 m, partly irregular

27

 

 

27a.  Subicular hyphae incrusted, spores spines up to 2 m, apiculus partly amyloid

 

         Tomentella lapida s.l.

 

         (incl. Tomentella fuliginea und Tomentella ramosissima)

 

27b.  Subicular hyphae not incrusted, spores with short spines

         Tomentella alpina Peintner & Dmmrich 2011

 

 

 

28a.  Spores with long spines up to 2,5 m, hymenophore smooth, fawn to rosy-buff,

 

         margin byssoid to fimbriat, rhizomorphs present

 

         Tomentella testaceogilva Bourd. & Galz. 1924

 

 

 

28b.  Not with the above combination of characters

29

 

 

29a.  Subhymenial hyphae to 8 (10) m wide, partly short celled and inflated

30

29b.  Subhymenial hyphae to 5,5 m wide, regular       

31

 

 

30a.  Spores rough tuberculate-echinulate 8-12 m,       

         a few warts bifurcate, mostly spores with oiltrops

         hyphae to 7 m wide, margin often fimbriat, partly rhizomorphs present

 

         Thelephora terrestris f. resupinata Donk 1933

 

         (Syn. ? Tomentella radiosa (P. Karst.) Rick 1934)

 

 

 

30b.  Spores with long spines 7-9 m,

         hyphae to 10 m wide, inflated up to 15 m wide,

 

         margin undetermined, rhizomorphs absent

 

         Tomentella sublilacina (Ellis & Holw.) Wakef. 1960

 

 

 

31a.  Subiculum partly thin, rhizomorphs rarely, 

         basidia partly with reddish contents (examines in distilled water!)

 

         hymenial with amber-tinted adhering material, solving in KOH

         and partly cyanescent 

32

 

 

31b.  Subiculum and rhizomorphs distinctly,    

 

         basidia not reddish in distilled water

34

 

 

32a.  Hymenophore brick-red, not cyanescent

        Tomentella lateritia Pat. 1894

 

 

32b. Hymenophore brownish

33

 

 

33a.  Hymeniophora dark brown, distinctly cyanescent            

 

         Tomentella molybdaea Bourdot & Galzin 1924   

 

33b.  Hymenophore pale brownish, not cyanescent

 

         Tomentella coerulea (Bres.) Hoehn. & Litsch. 1908

 

 

 

34a.  Basidia and hyphae cyanescent (hyphae and basidia partly bluish-green in KOH),

 

         hymenophore ferruginous-brown to olive-brown, granulose, 

 

         margin partly yellowish, rhizomorphs dimitic

 

         Tomentella ferruginea (Pers.) Pat. 1887

 

         (Remarks: If hymenophore blackish-brown view 17a.)

 

 

 

34b.  Basidia and hyphae not cyanescent

35

   

 

35a.  Hymenophore pale brownish with greenish or reddish tints, granulose, margin,

         byssoid, ferruginous-brown, rhizomorphs monomitic or rarely dimitic

 

         Tomentella punicea (Alb. & Schwein.) J. Schrt. 1889   

 

 

 

35b.  Hymenophore ferruginous-reddish to chestnut-brown, smooth to tubercular, 

 

         margin ferruginous-reddish, rhizomorphs dimitic

 

         Tomentella umbrinospora M.J. Larsen 1968

 

 

 

36a.  Spores ornaments bifurcate                                       

37

36b.  Spores ornaments not bifurcate                           

38

 

 

37a.  Hymenophore tubercular to hydnoid, cinnamon brown,

 

         spores slightly globose with short bifurcate ornaments 6-8 m,

 

         with thick-walled up to 200 m long tramal pseudocystidia

 

         Tomentella (Odontia) fibrosa (Berk. & Curtis) Kljalg 1996

 

 

 

37b.  Hymenophore smooth

 

         (cf. Pseudotomentella!)

 

 

 

38a.  Spores brown                                                            

38

38b.  Spores hyaline (cf. Tomentellopsis!)

 

 

 

39a.  Occasionally clamps present,

subicular hyphae and subhymenial hyphae occasionally clamps present,

spores regular to irregular ellipsoid, 8-12 m long, spines up to 1,5 m

Tomentella cinereoumbrina (Bres.) Stalpers 1993

 

 

 

39b.  All hyphae without clamps

40

 

 

40a.  Spores regular globose 79 m,  spines up to 3 m

 

         Tomentella spinosispora Cizek  2004

 

  (the type is described with clamps-bearing subicular hyphae,

but now recognised as alien hyphae, E. Martini 2021)

 

 

 

40b. Spores ellipsoid or irregular globose

41

 

 

41a.  Basidiocarps thick, crustose, margin usually byssoid to fimbriate, mostly on soil   

 

         Tomentella fuscocinerea (Pers.) Donk 1933

 

         (=Thelephora fuscocinerea Pers.)

 

 

 

41b.  Basidiocarps thin, tomentelloid, mostly on wood

42

 

 

42a.  Spores with spines up to 2,5 m, spores irregular globose,

 

         basidiocarps mucedinoid

 

         Tomentella badia (Link) Stalpers 1975

 

         (if spores regular globose view 42a.)

 

 

42b.  Spores with spines up to 1,5 m, spores regular ellipsoid

 

         Tomentella cf. brunneofirma M.J. Larsen 1967

 

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