Key of European species of Tomentella incl. Odontia 



-        spores size without ornaments

-        cyanescent-  hyphae and basidia bluish-green to bluish black in KOH






A- Hyphae clamped



B- Clamps absent or occasionally present






1a.   Cystidia present



1b.   Cystidia absent






2a.   Cystidia hyphoid 55-95 x 3-4 m, growing from subiculum and rhizomorphs,



        hymenophore hazel-brown to olive-brown, smooth to granulose,



        margin byssoid with brownish dimitic rhizomorphs,



        spores irregular 8,5-10 m 



        Tomentella atroarenicolor Nikol. 1970






2b.   Cystidia acuminate, capitate or clavate






3a.   Cystidia acuminate (basally widened, rarely hyphoid)



3b.   Cystidia capitate



3c.   Cystidia clavate






4a.   Hymenophore  hazel-brown, granulose,



        cystidia somewhat distended base and tapering towards the slightly,

        expanded apices, often apical encrusted with brown deposits,



        some spores reddish brown in KOH



        Tomentella subtestacea Bourd. & Galz.1924






4b.   Hymenophore greyish-green, smooth to granulose, mucedinoid,



        cystidia apical slightly incrusted



        Tomentella galzinii Bourd. 1924






5a.   Cystidia pale brownish to greenish



        basidiocarps separable, dimitic rhizomorphs present,



        hymenophore yellowish-brown to olive-brown,



        Tomentella pilosa (Burt) Bourd. & Galz. 1924






5b.   Cystidia hyaline,



        basidiocarps adherent, without rhizomorphs,



        hymenophore greyish-green to olive-brown,



        Tomentella viridula Bourd.& Galz.1924






6a.   Basidiocarps separable, rhizomorphs present,                             



        hymenophore greyish-brown to hazel-brown, smooth,
        subicular hyphae brownish, thick-walled


        Tomentella subpilosa Litsch 1960



        (similar to T. muricata, an American species)






6b.   Basidiocarps adherent, rhizomorphs absent,



        subicular hyphae hyaline and thin-walled                       





7a.   Spores irregular, partly lobed



        Tomentella clavigera Litsch. 1960






7b.   Spores regular globose to ellipsoid,



        subiculum thin to absent,



        subhymenial hyphae partly short celled and inflated



        Tomentella subclavigera Litsch. 1933






8a.   Spores average < 6,5m 



8b.   Spores average > 6,5 m






9a.   Spores 3-4 m,



        hymenophore with short spines, rusty yellowish



        Tomentella (Odontia) duemmeri (Wakef.) M.J. Larsen 1974






9b. Spores average 4,5-6,5 m 






10a.  Spores tubercular, hymenophore ferruginous-brown,



        smooth to hydnoid, margin byssoid, rhizomorphs dimitic



        Tomentella (Odontia) calcicola (Bourd. & Galz.) M.J. Larsen 1967





10b.  Spores echinulate






11a.  Basidiocarps adherent, rhizomorphs absent, hymenophore greyish brown



basidia up to 25 m long



Tomentella griseoumbrina Litsch. apud Lundell & Nannf. 1936  






11b. Basidiocarps separable, rhizomorphs present






12a. Hymenophore rosy brown, smooth



typical yellowish rhizomorphs present



Tomentella palliodocastanea X. Lu, Y.H. Mu & H.S. Yuan 2018






12b. Hymenophore greyish, granulose, rhizomorphs white



Tomentella cinerascens (P. Karst.) Hoehn. & Litsch. 1906






13a.  Spore ornaments bifurcate, Hymenophore hydnoid



13b.  Spore ornaments not bifurcate or with a view of bifurcate warts                              


Hymenophore smooth to granulare




14a.  Spores brown, slightly globose 7-8 m, with short pines,



         rhizomorphs dimitic



         Tomentella (Odontia) crinalis (Fr.) M.J. Larsen 1967        






14b.  Spores hyaline, irregular subglobose, 8-10 m


         rhizomorphs monomitic



         Tomentella italica (Sacc.) M.J. Larsen 1967




15a.  Immaturity basidia vesicular up to 25 m wide, 



         hymenophore smooth, fawn, basidiocarps compact,



         spores irregular subglobose 7-9,5 m



         Tomentella terrestris (Berk. & Broome) M.J. Larsen 1974






15b.  Immaturity basidia clavat to utriform, < 15 m wide                            






16a.  Spores average 6-7,5 m (if hyphae short celled and inflated



(if hyphae partly short celled and inflated go to 32a)



16b.  Spores average larger






17a.  Subicular hyphae thick-walled, brown, incrusted, up to 6 m wide,



         hymenophore greyish-brown to dark brown, granulose,



         spores partly irregular globose, 6-8 m


         Tomentella neobourdotii M.J. Larsen 1968





17b.  Subicular hyphae thin-walled, hyaline to pale brown 




18a.  Basidia and hyphae strong cyanescent, hymenophore blackish brown,



         rhizomorphs distinct, margin and subiculum ferruginous-yellowish



         Tomentella botryoides (Schwein.) Bourd. & Galz. 1924


(if rhizomorphs dimitic and hymenophore olivaceous brown



comp. T. ferruginea 36a.)




18b.  Basidia and hyphae not cyanescent, rhizomorphs absent



         hymenophore brownish, margin and subiculum same coloured, 



spores not bifurcate



         Tomentella sp.  FD 7548



(?Tomentella microspora (P. Karst.) Hhn. & Litsch. 1906 ss. Ales Jirsa)




19a. Hymenophore whitish, yellowish or pale ochraceous to mustard yellow

        Tomentella ochraceo-olivacea Litsch. 1933  

        (Syn. Tomentella fatrensis Svrček 1958)







19b. Hymenophor pale to dark brown, olivaceous or rusty brown






20a.  Spores subglobose or ellipsoid, not lobed          



20b.  Spores with irregular outline, mostly lobed     






21a.  Spores ellipsoid (lateral view)                     



21b.  Spores subglobose (lateral view)                        






22a.  Subicular hyphae hyaline to pale brownish, thin-walled,



         subhymenial hyphae short celled and inflated,



         spores 7-10 m long



         Tomentella ellisii (Sacc.) Jlich & Stalpers 1980






22b.  Subicular hyphae brown, thick-walled,



         subhymenial hyphae not short celled and inflated,



         spores 8-12 m long, partly irregular 



         Tomentella atramentaria Rostr. 1894




23a.  Subicular hyphae hyaline



23b.  Subicular hyphae brown






24a.  Hymenophore greyish-brown, tubercular, spores 8-11 m,



         basidiocarps separable, rhizomorphs present,



         subiculum white, margin yellowish to white



         Tomentella asperula (P. Karst.) Hoehn. & Litsch. 1906




24b.  Hymenophore pale brown to brick, spores 7-8,5 m



         (forms of T. lateritia 34a. and T. coerulea 35b. with more regular spores)





23a.  Hymenophor rust-yellowish, spores yellowish-brown



23b.  Hymenophor bright to dark brown or olive-greenish, spores brown




26a.  Spores with long spines, globose 8-11 m,

         apiculus amyloid

        Tomentella bryophila (Pers.) M.J. Larsen 1974




26b. Spores with short spines, irregular subglobose up to 6-9 m,

        apiculus not amyloid

        Tomentella ferruginella Bourd. & Galz. 1924




27a.  Hymenophore olive-greenish

         Tomentella viridescens (Bres. & Torr.) Bourd. & Galz. 1928 





27b.  Hymenophor bright to dark brown






28a.  Spores globose, 8 -11 m, spines up to 3 m,



         Subicular hyphae not incrusted



         Tomentella stuposa (Link) Stalpers 1984 s. l.

-- Spores 6 - 8 m Tomentella spec.

--      Spores 8 - 10 m  Tomentella stuposa (Link) Stalpers 1984 s. str.

--      Spores 10 - 13 m Tomentella bresadolae (Brinkmann) Hhn. & Litsch. 1908




28b.  Spores subglobose to ellipsoid, 7,5-9,5 m, partly irregular






29a.  Subicular hyphae incrusted, apiculus not amyloid, with spines up to 2 m



         Tomentella lapida (Pers.) Stalpers 1984



         (similar T. fuliginea with amyloid apiculus and subicular hyphae not incrusted)


29b.  Subicular hyphae not incrusted, with short spines, apiculus not amyloid

         Tomentella alpina Peintner & Dmmrich 2011



         (alpine species on Salix herbacea)






30a.  Spores with long spines up to 2,5 m, hymenophore smooth, fawn to rosy-buff,



         margin byssoid to fimbriat, rhizomorphs present



         Tomentella testaceogilva Bourd. & Galz. 1924






30b.  Not with the above combination of characters






31a.  Subhymenial hyphae to 8 (10) m wide, short celled and inflated



31b.  Subhymenial hyphae to 5,5 m wide, regular       






32a.  Spores rough tuberculate-echinulate,       

         a few warts bifurcate, mostly spores with oil drops, 

         margin often fimbriat, partly rhizomorphs present



         Tomentella radiosa(P. Karst.) Rick 1934



         (Syn. Thelephora terrestris f. resupinata Donk 1933)






32b.  Spores with long spines 7-9 m,

         hyphae to 10 m wide, inflated up to 15 m wide,



         margin undetermined, rhizomorphs absent



         Tomentella sublilacina agg.






33a.  Subiculum thin, rhizomorphs rarely, 

         basidia partly with reddish contents (examines in distilled water!)



         hymenial with amber-tinted adhering material, solving in KOH

         and partly cyanescent 






33b.  Subiculum and rhizomorphs distinctly,    



         basidia not reddish in distilled water






34a.  Hymenophore brick-red, not cyanescent

        Tomentella lateritia Pat. 1894





34b. Hymenophore brownish






35a.  Hymeniophore dark brown, distinctly cyanescent            



         Tomentella molybdaea Bourdot & Galzin 1924   


35b.  Hymenophore pale brownish, not cyanescent



         Tomentella coerulea (Bres.) Hoehn. & Litsch. 1908 ss. M.J. Larsen






36a.  Basidia and hyphae cyanescent,



         hymenophore ferruginous-brown to olive-brown, granulose, 



         margin partly yellowish, rhizomorphs dimitic



         Tomentella ferruginea (Pers.) Pat. 1887



         (Remarks: If hymenophore blackish-brown view Tomentella botryoides 19a.)






36b.  Basidia and hyphae not cyanescent






37a.  Hymenophore pale brownish with greenish or reddish tints, granulose, margin,

         byssoid, ferruginous-brown, rhizomorphs monomitic or rarely dimitic



         Tomentella punicea (Alb. & Schwein.) J. Schrt. 1889   






37b.  Hymenophore ferruginous-reddish to chestnut-brown, smooth to tubercular, 



         margin ferruginous-reddish, rhizomorphs dimitic



         Tomentella umbrinospora M.J. Larsen 1968






38a.  Spores ornaments bifurcate                                       



38b.  Spores ornaments not bifurcate                           






39a.  Hymenophore tubercular to hydnoid, cinnamon brown,



         spores slightly globose with short bifurcate ornaments 6-8 m,



         with thick-walled up to 200 m long tramal pseudocystidia



         Tomentella (Odontia) fibrosa (Berk. & Curtis) Kljalg 1996






39b.  Hymenophore smooth



         (cf. Pseudotomentella!)






40a.  Spores brown                                                            



40b.  Spores hyaline (cf. Tomentellopsis!)






41a.  Occasionally clamps present,


         subicular hyphae and subhymenial hyphae occasionally clamps present,

         spores regular to irregular ellipsoid, 8-12 m long, spines up to 1,5 m

         Tomentella cinereoumbrina (Bres.) Stalpers 1993






41b.  All hyphae without clamps






42a.  Spores regular globose 79 m,  spines up to 3 m



         Tomentella spinosispora Cizek  2004



         (the type is described with clamps-bearing subicular hyphae,

          but now recognised as alien hyphae, E. Martini 2021)






42b.  Spores ellipsoid or irregular globose






43a.  Basidiocarps thick, crustose, mostly on soil



         Tomentella fuscocinerea (Pers.) Donk 1933



         (= Thelephora fuscocinerea Pers.)






43b.  Basidiocarps thin, tomentelloid, on wood






44a.  Spores with spines up to 2,5 m, spores irregular globose,



         basidiocarps mucedinoid



         Tomentella badia (Link) Stalpers 1975



         (if spores regular globose view 42a.)




44b.  Spores with spines up to 1,5 m, spores regular ellipsoid



         Tomentella cf. brunneofirma M.J. Larsen 1967