Key
of European species of Tomentella incl. Odontia |
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-
spores
size without ornaments -
cyanescent-
hyphae and basidia bluish-green to bluish black in KOH |
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A- Hyphae clamped |
1 |
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B- Clamps absent or occasionally present |
38 |
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1a.
Cystidia present |
2 |
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1b.
Cystidia absent |
8 |
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2a.
Cystidia hyphoid 55-95 x 3-4 µm, growing from
subiculum and rhizomorphs, |
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hymenophore hazel-brown to olive-brown, smooth to granulose, |
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margin byssoid with brownish dimitic rhizomorphs, |
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spores irregular 8,5-10 µm |
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Tomentella atroarenicolor
Nikol. 1970 |
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2b.
Cystidia acuminate, capitate or clavate |
3 |
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3a.
Cystidia acuminate (basally widened, rarely hyphoid)
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4 |
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3b.
Cystidia
capitate |
5 |
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3c.
Cystidia
clavate |
6 |
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4a.
Hymenophore hazel-brown, granulose, |
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cystidia somewhat distended base and tapering towards the slightly,
expanded apices, often apical encrusted with brown deposits, |
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some spores reddish brown in KOH |
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Tomentella subtestacea
Bourd. & Galz.1924 |
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4b.
Hymenophore greyish-green, smooth to granulose, mucedinoid, |
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cystidia apical slightly incrusted |
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Tomentella galzinii
Bourd. 1924 |
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5a.
Cystidia pale brownish to greenish |
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basidiocarps separable, dimitic rhizomorphs present, |
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hymenophore yellowish-brown to olive-brown, |
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Tomentella pilosa
(Burt) Bourd. & Galz. 1924 |
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5b.
Cystidia
hyaline, |
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basidiocarps adherent, without rhizomorphs, |
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hymenophore greyish-green to olive-brown, |
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Tomentella viridula
Bourd.& Galz.1924 |
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6a. Basidiocarps
separable,
rhizomorphs present, |
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hymenophore greyish-brown to hazel-brown, smooth, |
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Tomentella subpilosa Litsch 1960 |
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(similar to T. muricata, an American
species) |
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6b.
Basidiocarps adherent, rhizomorphs absent, |
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subicular hyphae hyaline and
thin-walled
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7 |
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7a.
Spores irregular, partly lobed |
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Tomentella clavigera Litsch. 1960 |
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7b.
Spores regular globose to ellipsoid, |
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subiculum thin to absent, |
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subhymenial hyphae partly short celled and inflated |
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Tomentella subclavigera Litsch. 1933 |
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8a.
Spores average < 6,5µm |
9 |
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8b. Spores average > 6,5 µm |
13 |
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9a.
Spores 3-4 µm, |
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hymenophore with short spines, rusty yellowish |
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Tomentella (Odontia)
duemmeri (Wakef.)
M.J. Larsen 1974 |
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9b. Spores average 4,5-6,5 µm |
10 |
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10a. Spores
tubercular, hymenophore ferruginous-brown, |
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smooth to hydnoid, margin byssoid, rhizomorphs dimitic |
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Tomentella (Odontia)
calcicola (Bourd. & Galz.)
M.J. Larsen 1967 |
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10b. Spores
echinulate |
11 |
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11a. Basidiocarps adherent, rhizomorphs absent, hymenophore greyish brown |
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basidia up to 25 µm long |
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Tomentella griseoumbrina Litsch. apud Lundell & Nannf. 1936 |
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11b. Basidiocarps separable,
rhizomorphs
present |
12 |
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12a. Hymenophore rosy brown, smooth |
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typical yellowish rhizomorphs present
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Tomentella palliodocastanea X. Lu, Y.H. Mu
& H.S. Yuan 2018 |
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12b. Hymenophore
greyish, granulose, rhizomorphs white |
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Tomentella cinerascens (P. Karst.) Hoehn.
& Litsch. 1906 |
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13a. Spore ornaments bifurcate, Hymenophore hydnoid |
14 |
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13b.
Spore ornaments not bifurcate or with a view of bifurcate
warts |
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Hymenophore smooth to granulare |
15 |
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14a. Spores
brown, slightly globose 7-8 µm, with short pines, |
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rhizomorphs dimitic |
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Tomentella (Odontia) crinalis
(Fr.)
M.J. Larsen 1967
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14b. Spores
hyaline, irregular subglobose, 8-10 µm |
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rhizomorphs monomitic |
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Tomentella italica (Sacc.) M.J. Larsen 1967 |
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15a. Immaturity
basidia vesicular up to 25 µm wide, |
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hymenophore smooth, fawn, basidiocarps compact, |
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spores irregular subglobose 7-9,5 µm |
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Tomentella terrestris
(Berk. & Broome) M.J. Larsen 1974 |
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15b. Immaturity
basidia clavat to utriform, < 15 µm
wide
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16 |
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16a. Spores
average 6-7,5 µm (if hyphae short celled and inflated |
17 |
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(if hyphae partly short celled and
inflated go to 32a) |
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16b. Spores
average larger |
19 |
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17a. Subicular
hyphae thick-walled, brown, incrusted, up to 6 µm wide, |
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hymenophore greyish-brown to dark brown, granulose, |
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spores partly irregular globose, 6-8 µm |
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Tomentella neobourdotii M.J. Larsen 1968 |
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17b. Subicular
hyphae thin-walled, hyaline to pale brown |
18 |
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18a. Basidia
and hyphae strong cyanescent, hymenophore blackish brown, |
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rhizomorphs distinct, margin and subiculum ferruginous-yellowish |
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Tomentella botryoides (Schwein.) Bourd. & Galz. 1924 |
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(if rhizomorphs dimitic and hymenophore olivaceous brown |
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comp. T. ferruginea 36a.) |
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18b. Basidia
and hyphae not cyanescent, rhizomorphs absent |
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hymenophore brownish, margin and subiculum same coloured, |
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spores not bifurcate |
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Tomentella sp.
FD 7548 |
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(?Tomentella microspora (P.
Karst.) Höhn. & Litsch. 1906 ss. Ales Jirsa) |
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19b. Hymenophor pale to dark brown, olivaceous or rusty brown |
20 |
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20a. Spores subglobose or ellipsoid, not lobed |
21 |
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20b. Spores with irregular outline, mostly lobed |
30 |
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21a. Spores
ellipsoid (lateral
view) |
22 |
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21b. Spores
subglobose (lateral
view) |
23 |
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22a. Subicular
hyphae hyaline to pale brownish, thin-walled, |
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subhymenial hyphae short celled and inflated, |
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spores 7-10 µm long |
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Tomentella ellisii
(Sacc.) Jülich & Stalpers
1980 |
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22b. Subicular
hyphae brown, thick-walled, |
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subhymenial hyphae not short celled and inflated, |
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spores 8-12 µm long, partly irregular |
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Tomentella atramentaria
Rostr. 1894 |
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23a. Subicular
hyphae hyaline |
24 |
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23b. Subicular
hyphae brown |
25 |
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24a.
Hymenophore greyish-brown, tubercular, spores 8-11 µm, |
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basidiocarps separable, rhizomorphs
present, |
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subiculum white, margin yellowish to white |
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Tomentella asperula (P. Karst.) Hoehn. & Litsch. 1906 |
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24b.
Hymenophore pale brown to brick, spores 7-8,5 µm |
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(forms of T. lateritia 34a. and T. coerulea 35b. with more regular spores) |
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23a. Hymenophor
rust-yellowish, spores yellowish-brown |
26 |
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23b. Hymenophor
bright to dark brown or olive-greenish, spores brown |
27 |
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26a. Spores
with long spines, globose 8-11 µm,
apiculus amyloid Tomentella
bryophila (Pers.) M.J. Larsen 1974 |
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26b. Spores with
short spines, irregular subglobose up to 6-9 µm,
apiculus not amyloid
Tomentella ferruginella
Bourd. & Galz. 1924 |
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27a.
Hymenophore olive-greenish
Tomentella viridescens
(Bres. & Torr.) Bourd. & Galz. 1928 |
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27b. Hymenophor
bright to dark brown |
28 |
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28a. Spores
globose, 8 -11 µm, spines up to 3 µm, |
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Subicular hyphae not incrusted |
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Tomentella stuposa
(Link) Stalpers 1984 s. l. -- Spores 6 - 8 µm Tomentella
spec. --
Spores 8 - 10 µm Tomentella stuposa
(Link) Stalpers 1984 s. str. --
Spores 10 - 13 µm Tomentella bresadolae (Brinkmann) Höhn. & Litsch. 1908 |
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28b. Spores
subglobose to ellipsoid, 7,5-9,5 µm, partly irregular |
29 |
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29a. Subicular
hyphae incrusted, apiculus not amyloid, with spines up to 2 µm |
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Tomentella lapida (Pers.) Stalpers 1984 |
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(similar T. fuliginea with amyloid apiculus and subicular
hyphae not incrusted) 29b.
Subicular hyphae not incrusted, with short spines, apiculus
not amyloid
Tomentella alpina
Peintner & Dämmrich 2011 |
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(alpine species on Salix herbacea) |
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30a. Spores
with long spines up to 2,5 µm, hymenophore smooth, fawn to rosy-buff, |
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margin byssoid to fimbriat, rhizomorphs present |
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Tomentella testaceogilva
Bourd. & Galz. 1924 |
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30b. Not with
the above combination of characters |
31 |
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31a.
Subhymenial hyphae to 8 (10) µm wide, short celled and inflated |
32 |
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31b.
Subhymenial hyphae to 5,5 µm wide,
regular |
33 |
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32a. Spores
rough tuberculate-echinulate,
a few warts bifurcate, mostly spores with oil drops,
margin often fimbriat, partly rhizomorphs present |
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Tomentella radiosa(P. Karst.)
Rick 1934 |
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(Syn. Thelephora terrestris f. resupinata Donk 1933) |
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32b. Spores
with long spines 7-9 µm,
hyphae to 10 µm wide, inflated up to 15 µm wide, |
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margin undetermined, rhizomorphs absent |
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Tomentella sublilacina
agg. |
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33a. Subiculum
thin, rhizomorphs rarely,
basidia partly with reddish contents (examines in
distilled water!) |
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hymenial with amber-tinted adhering material,
solving in KOH
and partly cyanescent |
34 |
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33b. Subiculum
and rhizomorphs distinctly, |
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basidia not reddish in distilled water |
36 |
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34a.
Hymenophore brick-red, not cyanescent
Tomentella lateritia Pat.
1894 |
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34b. Hymenophore
brownish |
35 |
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35a. Hymeniophore dark brown, distinctly
cyanescent
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Tomentella molybdaea
Bourdot & Galzin
1924 35b. Hymenophore
pale brownish, not cyanescent |
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Tomentella coerulea
(Bres.) Hoehn.
& Litsch. 1908 ss. M.J. Larsen |
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36a. Basidia
and hyphae cyanescent, |
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hymenophore ferruginous-brown to olive-brown, granulose, |
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margin partly yellowish, rhizomorphs dimitic |
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Tomentella ferruginea (Pers.)
Pat. 1887 |
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(Remarks: If hymenophore blackish-brown view Tomentella botryoides 19a.) |
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36b. Basidia
and hyphae not cyanescent |
37 |
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37a.
Hymenophore pale brownish with greenish or reddish tints, granulose, margin,
byssoid, ferruginous-brown, rhizomorphs monomitic or rarely dimitic |
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Tomentella punicea (Alb. & Schwein.) J. Schröt. 1889 |
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37b.
Hymenophore ferruginous-reddish to chestnut-brown, smooth to
tubercular, |
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margin ferruginous-reddish, rhizomorphs
dimitic |
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Tomentella umbrinospora M.J. Larsen 1968 |
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38a. Spores
ornaments
bifurcate |
39 |
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38b. Spores ornaments
not
bifurcate |
40 |
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39a.
Hymenophore tubercular to hydnoid, cinnamon brown, |
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spores slightly globose with short bifurcate ornaments 6-8 µm, |
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with thick-walled up to 200 µm long tramal pseudocystidia |
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Tomentella (Odontia)
fibrosa (Berk. & Curtis) Köljalg 1996 |
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39b.
Hymenophore smooth |
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(cf. Pseudotomentella!) |
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40a. Spores
brown |
41 |
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40b. Spores hyaline (cf. Tomentellopsis!) |
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41a.
Occasionally clamps present, |
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subicular hyphae and subhymenial hyphae
occasionally clamps present,
spores regular to irregular ellipsoid, 8-12 µm long, spines up to 1,5 µm
Tomentella cinereoumbrina
(Bres.) Stalpers 1993 |
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41b. All hyphae
without clamps |
42 |
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42a. Spores
regular globose 7–9 µm, spines up to 3 µm |
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Tomentella spinosispora Cizek 2004 |
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(the type is described with clamps-bearing subicular hyphae,
but now recognised as alien hyphae,
E. Martini 2021) |
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42b.
Spores ellipsoid or irregular globose |
43 |
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43a.
Basidiocarps thick, crustose, mostly on soil |
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Tomentella fuscocinerea (Pers.) Donk 1933 |
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(= Thelephora fuscocinerea
Pers.) |
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43b.
Basidiocarps thin, tomentelloid, on wood |
44 |
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44a. Spores
with spines up to 2,5 µm, spores irregular globose, |
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basidiocarps mucedinoid |
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Tomentella badia
(Link) Stalpers 1975 |
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(if spores regular globose view 42a.) |
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44b. Spores
with spines up to 1,5 µm, spores regular ellipsoid |
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Tomentella cf. brunneofirma
M.J. Larsen 1967 |